I just read this in "Walking - the Complete Guide to the Complete Exercise" by Casey Myers:
In today's wild animal kingdom, running is used only as a survival gait. Dr Lovejoy said he doesn't know of any animal that runs more than 3 minutes at a time in its natural environment.
....... it is apparent that we are misusing the running gait when we employ it for prolonged periods of exercise.
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Huw, runnersdaily.co.uk